Sixteen lectotypifications of Asian Piper species are provided. Piper argyrites, P. baccatum, P. leptostachyum, P. majusculum, P. peepuloides, P. quinqueangulatum and P. sulcatum are accepted as species and many new synonyms are proposed. Useful diagnostic characters are described and geographical distribution data of each species are provided.
Fig 1
Type specimens of Piper species; (A) Piper argyrites Ridl. ex. C.DC, Ridley 8176 (K), (B) Piper baccatum Blume, Blume 624 (L), (C) Piper leptostachyum Wall. ex Miq., Wallich 6649 (K-W), (D) Piper majusculum Blume, Blume s.n. (L).
Fig 2
Figure 2. Type specimens of Piper species (A) Piper peepuloides Roxb., Wallich 6650A (U), (B-1) Chavica neesiana Miq., (B-2) Chavica sphaerostachya Miq., Wallich 6656 (U), (C) Piper quinqueangulatum Miq. Zollinger 1233 (P), (D) Piper sulcatum Blume, Blume s.n. (L).
Fig 3 P. argyrites
Figure 3. Piper argyrites Ridl. ex C.DC. (A) branch with an infructescence, (B) male inflorescence, (C) female inflorescence, (D) a portion of male inflorescence, (E) stamen, (F) floral bract (top view), (G) a portion of female inflorescence, (H) ovary (C. Suwanphakdee 246♂ & C. Suwanphakdee 247♀ (BK, BKF, KKU and QBG)). — Drawn by O. Kerdkaew.
Fig 4 P. baccatum
Figure 4. Piper baccatum Blume (A) branch with infructescences, (B) male inflorescence,
(C) female inflorescence, (D) a portion of male inflorescence, (E) male flower, (F) stamen, (G) a portion of female inflorescence, (H) ovary (C. Suwanphakdee 211♀ (BK, BKF and QBG), C. Suwanphakdee 285♂ (BKF)). — Drawn by O. Kerdkaew.
Fig 5 P. leptostachy
Figure 5. Piper leptostachyum Wall. ex Miq. (A) branch with a female inflorescence, (B) a portion of male inflorescence, (C) floral bracts (front and back views), (D) stamen, (E) infructescence (C. Suwanphakdee 464♀ (BK, BKF, KKU & QBG), C. Suwanphakdee 465♂ (BKF, KKU and QBG)). — Drawn by L. Loe-Khachon.
Fig 6 P. majusculum
Figure 6. Piper majusculum Blume (A) branch with an infructescence, (B) female flower, (C-D) floral bract (side & top views) (C. Suwanphakdee 54♀ (BK, BKF and KKU). — Drawn by P. Inthachub.
Fig 7 P. peepuloides
Figure 7. Piper peepuloides Roxb. (A) branch with an infructescences, (B) floral bract, (C) fruit (P. Srisanga 2249♀ (QBG)).— Drawn by O. Kerdkaew.
Fig 8 P. quinqueangulatum
Figure 8. Piper quinqueangulatum Miq. (A) branch with infructescences, (B) fruits & female flowers (C. Niyomdham 5332♀ (BKF and K)).— Drawn by O. Kerdkaew.
Fig 8 P. quinquean.tiff
Fig 9 P. sulcatum
Figure 9. Piper sulcatum Blume (A) branch with inflorescences,(B) a portion of female inflorescence, (C) ovary, (D) a portion of male inflorescence, (E) stamen, (F-G) floral bract (top & side views), (H) infructescences (C. Suwanphakdee 53♀ (BK, BKF and KKU) and C. Suwanphakdee 280♂ (BKFand QBG)). — Drawn by L. Loe-Khachon.
Fig 10
Figure 10. Piper argyrites Ridl. ex C.DC. (A) young leaves, (B) male inflorescence, (C) female inflorescence, (D) floral bract, (E) a portion of male inflorescence, (F) a portion of infructescence, (G) infructescence, Piper baccatum Blume (H) male inflorescence, (I) a portion of male inflorescence, (J) female flower, (K) male flower, (L) infructescence (photo by C. Suwanphakdee).
Fig 11
Figure 11. Piper leptostachyum Wall. ex Miq. (A) habit, (B) female inflorescence, (C) male inflorescence, (D) a portion of female inflorescence, (E) a portion of male inflorescence, (F) infructescences, (G) fruits, Piper majusculum Blume (H) branch with inflorescence, (I) a portion of infructescence (photo by C. Suwanphakdee).
Fig 12
Figure 12. Piper quinqueangulatum Miq. (A) habit, (B) male inflorescence, (C) female inflorescence, (D) infructescence, (E) fruits, Piper sulcatum Blume (F) female inflorescence, (G) a portion of female inflorescence, (H) infructescence (photo by C. Suwanphakdee).
