Data from: Does evenness even exist?
Data files
Jul 10, 2023 version files 1.86 MB
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Ecological_Register_data.txt.gz
1.53 MB
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Ecological_Register_references.txt.gz
201.80 KB
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Ecological_Register_samples.txt.gz
126.76 KB
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README.md
1.07 KB
Nov 19, 2023 version files 1.86 MB
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Ecological_Register_data.txt.gz
1.53 MB
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Ecological_Register_references.txt.gz
201.80 KB
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Ecological_Register_samples.txt.gz
126.76 KB
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README.md
1.23 KB
Mar 20, 2024 version files 1.86 MB
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Ecological_Register_data.txt.gz
1.53 MB
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Ecological_Register_references.txt.gz
201.80 KB
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Ecological_Register_samples.txt.gz
126.76 KB
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README.md
1.22 KB
Jul 03, 2025 version files 1.86 MB
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Ecological_Register_data.txt.gz
1.53 MB
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Ecological_Register_references.txt.gz
201.80 KB
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Ecological_Register_samples.txt.gz
126.76 KB
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README.md
1.54 KB
Abstract
The idea that diversity is a combination of species richness and the so-called "evenness" of count distributions is a bedrock concept in ecology. Researchers often compute stand-alone evenness indices. They also examine Hill numbers related to Shannon's H and Simpson's D because these metrics balance richness and "evenness" to various degrees. But evenness is an operationally problematic abstraction, not a thing out in the world. Evenness indices and Hill numbers in empirical data are overly sensitive to the abundance of dominant species, poorly replicable within communities, highly variable among similar communities, and a weak indicator of latitudinal biodiversity trends. They are inconsistently related to the parameters of key models that might underlie count distributions, and they vary highly in simulation even when these model parameters do not vary. Ecologists would benefit by instead determining which real distributions fit which theoretical models and using estimated parameters to understand community structure and assembly.
This file includes all species inventories downloaded from the Ecological Register on 23 October 2022.
Description of the data and file structure
Ecological_Register_data.txt.gz is a single self-contained, tab delimited data matrix including a header that describes the data columns. The file is a list of species occurrences within species inventories. The reference.no field points to a list of references to publications in the Ecological_Register_references.txt.gz file. The sample no field points to a table with additional species inventory metadata to be found in the Ecological_Register_samples.txt.gz file. The blank cells in the files represent cases where no data were entered into the relevant fields, and will be interpreted as NAs when uploaded by an R script. There are no hidden values.
Sharing/Access information
The Ecological Register website is at http://ecoregister.org.
Code/Software
Code used in the paper “Does evenness even exist?” is reposited on Zenodo (https://doi.org/10.5281/zenodo.15762247). The underlying code used to run the analyses of the preprint “Three models of ecological community assembly” is in the richness.tar.gz file that includes the R library called richness. The current version of the richness package is at https://github.com/johnalroy/richness/releases. The code used to prepare the figures, tables, and individual statistics reported in the text of “Three models” is included in the script called three_models.R.
The gzipped Ecological Register data file (Ecological_Register_data.txt.gz) is a full set of published species inventories of trees and terrestrial animals downloaded from the Ecological Register website on 23 October 2022. The additional files include metadata pertaining to the references used to document the inventories (Ecological_Register_references.txt.gz) and to the species inventories themselves (Ecological_Register_samples.txt.gz) .The gzipped and tarred richness R library (richness.tar.gz) was used to prepare the data and analyses in a preprint of an earlier paper called "Three models of ecological community assembly". The blank cells in the .txt files represent cases where no data were entered into the relevant fields, and will be interpreted as NAs when uploaded by an R script. There are no hidden values.
The R programming environment is required to open the richness library and the scripts.