1. Re: LambkinWorldBombs2003.nex 2. Introduction o This NEXUS file, built using the program Mesquite version 2.74 (Maddison, WP and Maddison, DR, 2010) contains the scoring of 78 beeflies (Diptera: Bombyliidae: Anthracinae) from the World fauna of Exoprosopini for 207 morphological characters as reported in Lambkin CL, Yeates DK, and Greathead DJ (2003) An evolutionary radiation of bee flies in semi-arid Australia: Systematics of the Exoprosopini (Diptera: Bombyliidae). Invertebrate Systematics 17: 735-891. o Note also contains and one most parsimonious tree (one of four) generated using PAUP*4.0b10 (Swofford, DL, 2002) as reported in Lambkin CL, Yeates DK, and Greathead DJ (2003) An evolutionary radiation of bee flies in semi-arid Australia: Systematics of the Exoprosopini (Diptera: Bombyliidae). Invertebrate Systematics 17: 735-891. o Data collation by Chris Lambkin o Contact for information: Dr Christine L Lambkin: Entomology, Queensland Museum, PO Box 3300, South Brisbane, Queensland, Australia 4101 (christine.lambkin@qm.qld.gov.au). 3. Description of file format: o column headings The taxon list is given below: 1 Comptosia 2 Docidomyia 3 Petrorossia 4 Brachyanax 5 Anthrax_maculatus 6 Villa_fuscicostata 7 Lepidanthrax 8 Diatropomma 9 Exoprosopa_pandora 10 Exoprosopa_batrachoides 11 Exoprosopa_capucina 12 Exoprosopa_cleomene 13 Exoprosopa_dux 14 Exoprosopa_heros 15 Exoprosopa_jacchus 16 Exoprosopa_rivularis 17 Exoprosopa_sylvana 18 Atrichochira_commoni 19 Atrichochira_inermis 20 Atrichochira_paramonovi 21 Atrichochira_pediformis 22 Collosoptera 23 Exoprosopa_aurivestrix 24 Defilippia_busiris 25 Defilippia_eritreae 26 Defilippia_efflatouni 27 Defilippia_gentilis 28 Defilippia_luteicosta 29 Defilippia_minos 30 Defilippia_nigrifimbriata 31 Defilippia_rubescens 32 Exoprosopa_villaeformis 33 Euligyra_enderleini 34 Euligyra_mars 35 Euligyra_monacha 36 Euligyra_paris 37 Euligyra_sisyphus 38 Heteralonia_daveyi 39 Heteralonia_dubia 40 Heteralonia_lepidogastra 41 Heteralonia_oculata 42 Heteralonia_megerlei 43 Heteralonia_melanoptera 44 Heteralonia_mucorea 45 Heteralonia_pygmalion 46 Heteralonia_singularis 47 Heteralonia_suffusa 48 Heteralonia_umbrosa 49 Hyperalonia 50 Ligyra_bombyliformis 51 Ligyra_punctipennis 52 Ligyra_satyrus 53 Ligyra_sinuatifascia 54 Litorhina_dentifera 55 Litorhina_erythraeus 56 Exoprosopa_inaequalipes 57 Litorhina_macroptera 58 Litorhina_pseudocollaris 59 Exoprosopa_punctulata 60 Micomitra_parvicellula 61 Micomitra_stupida 62 Pseudopenthes_fenestrata 63 Pseudopenthes_hesperis 64 Pterobates_apicalis 65 Pterobates_pennipes 66 Balaana_efflatounbeyi 67 Balaana_gigantea 68 Balaana_grandis 69 Balaana_latelimbata 70 Balaana_obliquebifasciata 71 Balaana_onusta 72 Balaana_tamerlan 73 Kapu_adelaidica 74 Munjua_trigona 75 Muwarna_stellifera 76 Palirika_marginicollis 77 Palirika_decora 78 Wurda_windorah The character list that forms columns 2-208 is as follows: For a full description of these characters see Appendix 2 in Lambkin CL, Yeates DK, and Greathead DJ (2003) An evolutionary radiation of bee flies in semi-arid Australia: Systematics of the Exoprosopini (Diptera: Bombyliidae). Invertebrate Systematics 17: 735-891. Morphological terminology follows that of McAlpine (1981), Yeates (1994), and Lambkin et al. (2003). MORPHOLOGICAL CHARACTERS HEAD Frontal 1. Head scales: (0) absent; (1) present 2. Face scale density: (0) absent; (1) sparse; (2) overlapping; (3) carpet 3. Frons scale density: (0) absent; (1) sparse; (2) overlapping; (3) carpet; (4) distinct dense medial patch 4. Frons with vertical groove: (0) absent; (1) depression; (2) groove 5. Frons with horizontal depression: (0) absent; (1) shallow; (2) distinct; (3) deep Dorsally 6. W head/W thorax: (0) <; (1) =; (2) > 7. L antennae to compound eye/L scape: (0) <; (1) > or =; (2) > or = 2x; (3) > or = 4x 8. L antennal separation/L scape: (0) <; (1) > or =; (2) > or = 2x; (3) > or = 3x 9. Male compound eye separation /W OT: (0) meet; (1) <; (2) =; (3) < or = 2x; (4) < or = 3x; (5) > 3x 10. Female compound eye separation /W OT: (0) < or = 2x; (1) < or = 3x; (2) > 3x 11. OT to posterior margin of compound eye/ L OT: (0) < or = OT; (1) > OT; (2) > or = 2x; (3) > or = 3x 12. L occiput/L OT: (0) < or = 2x; (1) < 3x; (2) occiput long, well developed > or = 3x 13. L vertex; i.e. L OT to occipital groove/ L OT: (0) < or = L OT; (1) < or = 2x; (2) wide > 2x 14. Depth occipital foveal depression: (0) no vertex; (1) not depressed; (2) shallow; (3) deep, slopes posteriorly at 45 degrees to a short OG 15. W occipital foveal depression: (0) no vertex; (1) not depressed; (2) narrower than compound eye separation; (3) wider than compound eye separation 16. Apical occipital groove: (0) narrow; (1) wide rounded Lateral 17. Shape of head laterally: (0) round; (1) protruding but rounded, blunt; (2) conical 18. L proboscis: (0) < L oral cavity; (1) > L oral cavity; (2) > L head; (3) > 1.5x L head; (4) > 2x L head 19. L palps/ L proboscis: (0) rudimentary; (1) < 0.25x, short; (2) < 1, long 20. Genae setae surrounding oral cavity: (0) not grouped; (1) grouped laterally; (2) grouped apically, small tuft 21. L setae below antennae/L scape: (0) >; (1) < or =; (2) < or = 0.5 22. Horizontal depression between antennae: (0) absent or shallow; (1) distinct; (2) deep 23. W of face projection/W compound eye including indentation posterior margin of eye: (0) < 1/4; (1) < or = 1/2; (2) < or = 1 24. W of the indentation on the posterior margin of the compound eye/L OT: (0) < or = L OT; (1) > L OT; (2) > or = 2x L OT 25. L of the line from the posterior margin of compound eye bisecting the compound eye facets/L OT: (0) absent; (1) <; (2) > or =; (3) > or = 2x ANTENNAE 26. L scape /L pedicel: (0) < or =; (1) < or = 3x; (2) > 3x 27. L PP/ L pedicel: (0) < or = 3x; (1) < or = 4x; (2) > 4x 28. PP base shape: (0) broad not round base; (1) onion-like, round base abruptly narrowed; (2) medially divided, laterally; (3) conical, broad base, gradually narrowed 29. L PP rod/L base: (0) long > 2x; (1) short < 2x; (2) no thin rod 30. Distinct joint between PP and BSM: (0) absent; (1) present 31. L BSM/L pedicel: (0) absent; (1) < or =; (2) < or = 2x; (3) < or = 3x; (4) > 3x 32. BSM apical hairs: (0) absent; (1) present 33. L ASM/W BSM: (0) < W BSM, minute spine; (1) > W BSM; (2) > 2x; (3) conical twisted hat THORAX 34. Collar Ma: (0) pointed; (1) midstyle; (2) pectinate 35. Scm reflective vestiture: (0) bright; (1) very dull dark; (2) not reflective 36. Ma AN: (0) pointed; (1) midstyle; (2) pectinate 37. L PR bristles/ L PN: (0) > 2x; (1) >; (2) <; (3) absent 38. LT vestiture: (0) bare; (1) some hair; (2) dense hair 39. Ma LT: (0) absent; (1) pointed; (2) midstyle; (3) pectinate 40. MT vestiture: (0) bare; (1) some hair; (2) dense hair 41. L PA bristles/Scu: (0) absent; (1) < 0.5; (2) < or =; (3) > 42. L thorax/Scu: (0) < or = 2x; (1) < or = 3x; (2) > 3x 43. Scu vestiture reflective: (0) bright; (1) very dull dark; (2) absent Legs 44. C1 very long setae: (0) absent; (1) some; (2) dense 45. L forefemur/ L coxa: (0) < or = 1.5x; (1) < or = 2x; (2) < or = 2.5x; (3) > 2.5x 46. Forefemoral spines: (0) absent; (1) short < W femur; (2) long 47. Forefemoral long hairs: (0) absent; (1) some; (2) dense 48. Foretibial spicules: (0) absent; (1) some; (2) dense 49. L foretarsus/ L foretibia: (0) > or = 1; (1) > or = 0.75; (2) > 0.5 50. Foretarsal microchaetae: (0) absent; (1) very few < 10; (2) present 51. Foretarsal microchaetae: (0) absent; (1) ends bulbous; (2) ends slightly bent; (3) ends distinctly bent 52. L foreclaw/ L midclaw: (0) < claw; (1) < or = half; (2) < or = third 53. Midfemoral spines: (0) absent; (1) short < W femur; (2) some long 54. Midfemoral long hairs: (0) absent; (1) some; (2) dense 55. Midtibial spicules: (0) absent; (1) some; (2) dense 56. L midpulvilli/ L claw: (0) > or =; (1) <; (2) < or = half; (3) < 0.2 57. Midpulvilli: (0) large, flattened, membranous; (1) small rounded setose; (2) chisel-conical 58. Hindfemoral spines: (0) absent; (1) short < 0.5 W femur; (2) some long 59. Hindfemur long hairs: (0) absent; (1) some; (2) dense 60. Long hindtibial scales: (0) no long scales; (1) some long scales; (2) fluffy - protruding; (3) feathery; (4) very long, dense, feathered fringes 61. Hindtibial spicules: (0) absent; (1) some spicules; (2) apical patch; (3) dense spicules 62. Hindtibial spicules L: (0) absent; (1) same; (2) inner row longer 63. L hindpulvilli/L claw: (0) > or =; (1) <; (2) < or = half; (3) < or = 0.2 64. Hindpulvilli: (0) large, flattened, membranous; (1) small rounded setose; (2) chisel -conical Wing 65. Patagium: (0) absent hairs only; (1) present scales 66. Basicosta: (0) absent; (1) blunt; (2) sharp Wing venation 67. Crossvein forming an extra anterior apical submarginal cell: (0) absent; (1) extra apical submarginal cell 68. R2+3 join R4+5: (0) basal to r-m > L r-m; (1) at r-m 69. R2+3 rises from R4+5: (0) acutely; (1) at right angles 70. 2nd r-m crossvein: (0) absent; (1) present 71. Spurvein base R2+3: (0) absent; (1) bump or present 72. R2+3 apical loop: (0) long apical loop; (1) loop > 180 degrees; (2) at least 90 degrees bend; (3) absent 73. i-r1, R2+3 to R4: (0) absent; (1) present 74. i-r1 crossvein: (0) absent; (1) straight; (2) slightly sinuous; (3) distinctly sinuous 75. Spurvein i-r: (0) absent; (1) bump or present 76. L i-r1/L r-m: (0) absent; (1) < or =; (2) < or = 2x; (3) < 3x; (4) > or = 3x 77. Spur-vein base R4+5: (0) absent; (1) bump or present 78. Spur-vein R4: (0) absent; (1) bump or present 79. R5/R1 meet wing: (0) R5 distal to R1; (1) equal; (2) R5 basal to R1 80. i-r2, R4 to R5: (0) absent; (1) present 81. M1: (0) straight; (1) slightly sinuous; (2) sinuous 82. L m-m/r-m: (0) < or = 2x; (1) < or = 3x; (2) > 3x 83. m-m: (0) straight; (1) slightly sinuous; (2) sinuous 84. m-m spurvein: (0) absent; (1) into m2; (2) into discal; (3) crossvein form basal cell 85. m-m to hind wing margin: (0) oblique; (1) parallel; (2) horizontal 86. M2: (0) straight; (1) slightly sinuous; (2) sinuous 87. r5 open: (0) open; (1) narrow < r-m; (2) just closed; (3) closed and stalked - acute; (4) closed and stalked -obtuse 88. m1 open: (0) open; (1) closed and stalked 89. m2 open: (0) open; (1) closed and stalked - acute 90. L mcu/r-m: (0) > 3x; (1) < 3x; (2) < or = 2x; (3) < or = 91. m-cu: (0) straight; (1) slightly sinuous; (2) 90 degrees basally; (3) sinuous 92. Spur-vein m-cu: (0) absent; (1) spur into discal cell 93. Spur-vein into M2: (0) absent; (1) spur-vein into m2; (2) cross-vein to CuA1 94. W anal/ W posterior cubital: (0) < or =; (1) < or = 1.5x; (2) < or = 2x; (3) > 2x 95. cup open: (0) open; (1) narrow < r-m; (2) closed at wing margin (Fig. 9B); (3) closed and stalked - acute 96. Anal lobe margin: (0) hairs; (1) some scales; (2) scales dense 97. Anal cell: (0) broad rounded; (1) rounded; (2) thin linear; (3) very reduced 98. Alula reduced: (0) not reduced; (1) reduced, L < 4x W 99. Alula margin: (0) hairs; (1) some scales; (2) scales dense 100. Squamal margin: (0) hairs; (1) some scales; (2) scales dense 101. Squama reduced: (0) not reduced; (1) reduced, L < 4x W 102. wing L: (0) < or = 10; (1) 10-15; (2) 16-20; (3) 21-25; (4) > 25 103. wing L/W: (0) < or = 3x; (1) long > 3x 104. L wing/ L abdomen: (0) > or = 3x; (1) > or = 2 x; (2) > 1.5 x; (3) > ABDOMEN 105. Abdomen apically: (0) rounded; (1) narrowed; (2) truncate, parallel sided 106. Abdomen L/ W T2: (0) > 2.5x; (1) < 2.5x; (2) < 2x; (3) < or = 1.5x; (4) < or = 107. Abdominal vestiture reflective: (0) bright; (1) very dull dark; (2) not reflective 108. Abdominal bristles/hair: (0) dorsally and laterally; (1) lateral and apically T7; (2) apically; (3) absent 109. Long lateral hairs > T1: (0) dense tufts; (1) some; (2) absent 110. Ma T1: (0) pointed; (1) midstyle; (2) pectinate 111. Scales: (0) absent; (1) adpressed scales; (2) upstanding scales; (3) long upstanding scale tufts Male genitalia 112. Male genitalia twisted: (0) no twisting, gonocoxae ventral; (1) 90 degrees; (2) 180 degrees, gonocoxae dorsal 113. E setae grouping: (0) not grouped; (1) medioapically; (2) lateroapically; (3) laterally 114. E setae group: (0) not grouped; (1) loose; (2) strong; (3) dense tufts 115. Epandrial spines: (0) absent; (1) short and broad; (2) long 116. E apically: (0) deeply indented; (1) concave-indented; (2) truncate; (3) convex rounded; (4) convex pointed 117. E apical flange: (0) absent; (1) < quarter base; (2) < third base; (3) < half base; (4) > half rest base 118. E medial flange: (0) absent; (1) slight; (2) distinct < quarter base 119. L E basal flange: (0) absent; (1) < quarter base; (2) < third base (Fig. 10B); (3) < half base; (4) > or = half rest base; (5) > base 120. Mid W/L basal flange: (0) absent; (1) < quarter length base; (2) < half length; (3) > half length (4) > length 121. E posterolateral flange: (0) absent; (1) < quarter length base 122. E basal flange recurved: (0) absent (Fig. 10B); (1) < quarter base; (2) < third base; (3) < half base; (4) > half rest base; (5) > base 123. E basally extended: (0) absent; (1) present 124. SES: (0) absent; (1) linear; (2) triangular; (3) quadrate; (4) single (Fig. 10A) 125. L SES/G W: (0) absent; (1) < eighth; (2) < quarter; (3) > quarter 126. G setae: (0) some; (1) dense; (2) tufts 127. G setae group: (0) absent; (1) not grouped; (2) apically; (3) medially; (4) laterally; (5) basally 128. Thick G setae number: (0) absent; (1) no thick setae; (2) some; (3) many; (4) 6-8 long 129. Thick G setae position: (0) absent; (1) no thick setae; (2) apically; (3) medially; (4) laterally; (5) basally 130. G subapical indentation: (0) absent; (1) slight < third; (2) narrowed apically > third 131. W G medial indentation: (0) absent; (1) < third; (2) > third; (3) > half 132. G medial weakness: (0) absent; (1) desclerotised line; (2) lines of weakness; (3) division medially 133. G ventral division: (0) line fusion basally; (1) line fusion entire; (2) fused medially; (3) fused basally; (4) fused 134. G medioventrally: (0) deeply indented; (1) indented medially; (2) flat; (3) convex shell 135. G ventral ridge: (0) absent; (1) slight; (2) distinct 136. G basal projection of the ventral ridge: (0) absent; (1) slight; (2) distinct; (3) recurved hook 137. G basomedial margin: (0) deeply indented; (1) indented, concave; (2) smooth, linear; (3) convex 138. H: (0) absent; (1) present 139. H laterally: (0) absent; (1) indented between G, smooth; (2) projecting; (3) with spur; (4) with finger (Fig. 10F) 140. RM: (0) small; (1) > L GS; (2) large recurved 141. L G A: (0) absent; (1) < GS; (2) > GS 142. G plates dorsoapically: (0) medially parallel; (1) angled basomedial plates diverge dorsally 143. G dorsoapical plates extension apically: (0) not extended apically; (1) small apical extension; (2) long apical extension beyond the base of the gonostyli 144. GS: (0) large base, long pointed flange; (1) laterally bifid; (2) simple curved hook; (3) medial hook; (4) lateral hook 145. GS basal projection: (0) absent; (1) small; (2) < GS 146. L AE: (0) < GS; (1) = GS; (2) > GS 147. AE EP separate: (0) absent; (1) present 148. EP: (0) absent; (1) present 149. EP deep ventral notch: (0) no EP; (1) absent; (2) medial 150. EP expanded apically: (0) no EP; (1) not expanded (Fig. 10H); (2) < or = 2x neck; (3) < 3x neck; (4) > 3 x neck 151. EP apical plate: (0) no EP; (1) absent; (2) apical plate 152. EP medioventral projection: (0) no EP; (1) absent; (2) above AE 153. EP lateroapical lobes: (0) no E; (1) absent; (2) rounded dorsally; (3) pointed dorsally 154. EP lateral projection laterally: (0) no EP; (1) absent (2) lateral 155. EP medial projection laterally: (0) no EP; (1) absent; (2) medial 156. EP pair ventral projections: (0) no EP; (1) absent; (2) medial below AE 157. L EP: (0) no EP; (1) < GS base; (2) > GS base; (3) > G margin 158. EP recurved apicomedial projection: (0) no EP; (1) absent; (2) present dorsally 159. EP recurved apically: (0) no E; (1) absent; (2) apex recurved lateral view 160. EP apical setae: (0) no EP; (1) absent; (2) present 161. BP expanded: (0) not expanded; (1) round; (2) swollen spherical; (3) bilobed 162. LAEA: (0) spoon convex up; (1) spoon concave; (2) linear; (3) absent 163. L lateral AE A: (0) < L GS; (1) < G margin; (2) = G margin; (3) absent 164. AAES: (0) spoon convex; (1) spoon concave; (2) narrow wedge; (3) linear 165. AAES: (0) < B; (1) = L GS; (2) < G margin; (3) to G margin 166. EJA: (0) racquet - round; (1) linear 167. L EJA: (0) within G; (1) = G; (2) > G < L GS; (3) > G > L GS Female genitalia 168. AC spines: (0) 3 prs; (1) 4 prs; (2) 5 prs; (3) > 5 prs; (4) > 10 prs 169. AC spines apically: (0) thin tapering; (1) broader spoon shaped 170. T9+10 sclerites: (0) 1 sclerite; (1) 3 sclerites 171. T9 dorsal medioapical unsclerotised lacuna: (0) absent; (1) present 172. T8 dorsal medioapical unsclerotised lacuna: (0) absent; (1) present 173. T8 hair: (0) apical half; (1) apical half bare medially; (2) apical edge 174. T8 laterally: (0) not indented; (1) indented arms 175. T8 A divided medially: (0) not divided; (1) slightly; (2) distinctly > or = half width 176. T8 A lateral projections: (0) absent; (1) slight; (2) not linear; (3) linear 177. T8 A L/medial W: (0) margin thickened, sclerotised; (1) < or = quarter; (2) < or = half; (3) <; (4) > or =; (5) > 2x; (6) > or = 3x 178. T8 A internal structure: (0) absent; (1) linear; (2) quadrate plate 179. S8 sclerites: (0) 1 linear 2 round; (1) 1 linear 2 round 1 medial; (2) U-shaped 2 triangular; (3) 2 round; (4) sheet 180. Furca: (0) U-shaped; (1) 3 separate sclerites; (2) 2 rods 181. ST tube: (0) short < third L pump; (1) present; (2) long > x 8 L pump 182. Basal endplate: (0) absent; (1) small; (2) present (Fig. 9H); (3) large 183. Basal endplate: (0) absent; (1) simple-thin processes; (2) thick processes; (3) funnel 184. Sperm pump: (0) short pump; (1) very long pump 185. Long pump papillae: (0) no long papillae; (1) unpigmented; (2) pigmented 186. Pump processes basally: (0) no processes; (1) short processes; (2) long papillae 187. Pump processes medially: (0) no processes; (1) short processes; (2) long papillae 188. Pump processes apically: (0) no processes; (1) short processes 189. Apical endplate: (0) large; (1) present; (2) small; (3) absent 190. Apical endplate: (0) absent; (1) simple- thin processes; (2) thick processes; (3) funnel; (4) double 191. SR: (0) L < or = W; (1) L > W; (2) L > 2 W; (3) L > 4 W; (4) L > 6 W; (5) L > 8 W; (6) L > 30 W 192. ST basal sclerotised plate: (0) absent; (1) basal sclerotised plate 193. Tube ST to pump: (0) absent; (1) present; (2) long 194. SR shape: (0) round square; (1) oval; (2) pear, expanded apically; (3) long; (4) expanded basally 195. SR apically: (0) rounded blunt; (1) nipple; (2) narrowed; (3) knob 196. ST round basal bulb: (0) no round BB; (1) round BB 197. SR pigmented: (0) unpigmented; (1) pigmented; (2) basally unpigmented 198. ST long medial tube: (0) no medial tube; (1) tube medially 199. Long membranous base: (0) no long base; (1) long MB 200. ST long MB basally swollen: (0) no long base; (1) symmetrically; (2) asymmetrically 201. ST clear rings: (0) no rings; (1) clear ring; (2) long striated collar 202. SR to pump: (0) symmetrical; (1) asymmetrical 203. SR medially bent: (0) absent; (1) bent reservoir 204. SR apically bent: (0) absent; (1) tip only 205. Tubules: (0) absent; (1) present 206. SR walls: (0) thin unsclerotised; (1) thick sclerotised 207. SR walls: (0) no dimples; (1) with dimples thin unsclerotised o missing data = ?, inapplicables/gap = - o abbreviations used see below: Figure numbers refer to CL Lambkin and JS Bartlett ZooKeys 2011 Bush Blitz aids description of three new species and a new genus of Australian beeflies (Diptera: Bombyliidae: Exoprosopini) Morphological Abbreviations A apodeme (Fig. 6F) AAES anterior arms of aedeagal sheath (Fig. 5E) AC acanthophorite (Fig. 6F) AE aedeagus (Fig. 5F-H) AG accessory glands (Fig. 9H) AN anepisternum (mesopleuron) APA anterior postalar ridge ASM antennal apical stylomere BSM basal stylomere BB spermathecal basal bulb BP basiphallus (Fig. 5E, F) BSM antennal basal stylomere C costa Cu cubital vein dc discal cell E epandrium EJA ejaculatory apodeme (Fig. 5E, F) EP epiphallus (Fig. 5E, F) F flagellomere G gonocoxa GS gonostylus (Fig. 5B) H hypandrium (Fig. 5C, D) i-r1 wing inter-radial crossvein between veins R2+3 and R4; i-r2 wing inter-radial crossvein between veins R4 and R5. K katepisternum (sternopleuron) L length LAEA lateral aedeagal apodemes (Fig. 5E, F) LT laterotergite (metapleuron) M medial vein Ma macrochaetae MB membranous base mcu section of wing vein CuA1 between vein M3 and crossvein m-cu m-m basal section of wing vein M2 MT mediotergite (metanotum) OG occipital groove OT ocellar triangle PA postalar PE proepimeron (prosternum) PL plumula PN postpronotal lobe (humeral callus) PP postpedicel PR prealar R radial vein 1st r2+3 wing cell r2+3 basal to crossvein i-r1 2nd r2+3 wing cell r2+3 apical to crossvein i-r1 RM ramus (Fig. 5F, H) S sternite (Fig. 6F, G) SES subepandrial sclerites (Fig. 4E) Sc subcostal vein Scm scutum Scu scutellum SP sperm pump (Fig. 9H) SR spermathecal reservoir (Fig. 9H) ST spermathecal SS scutoscutellar suture T tergite (Fig. 6F, G) TR tympanal ridge W width WR wing root