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Supplements for: Review and revision of the Olivoidea (Neogastropoda) from the Paleocene and Eocene of the U.S. Gulf Coastal Plain


Allmon, Warren; Friend, Dana (2022), Supplements for: Review and revision of the Olivoidea (Neogastropoda) from the Paleocene and Eocene of the U.S. Gulf Coastal Plain, Dryad, Dataset,


Numerous species of “oliviform” gastropods have been recognized in the Paleogene of the U.S. Gulf Coastal Plain, many of which have previously been allied to the “Bullia group” in the family Nassariidae, and placed in a variety of poorly-defined genera. We review these species, revise their generic and familial placement, and present a phylogenetic analysis. Of 19 species considered valid, all are assigned to Olivoidea, six to Olividae -- one to Oliva, five to Agaronia -- and the rest to Ancillariidae. The highly variable species Ancillaria altile Conrad, 1832 is referred in the genus Ancillopsis and appears to have evolved anagenetically over an interval of more than 15 million years. Ancillaria tenera Conrad, 1834a and Ancillaria scamba Conrad, 1832 are placed in the new genus Palmoliva. Monoptygma Lea, 1833 is demonstrated to belong to Ancillariidae, and to contain only a single species. Some specimens assigned to Lisbonia expansa Palmer, 1937 are split into adults assigned to Ancillopsis altilis and juveniles (together with several other species) in Anbullina elliptica (Whitfield, 1865). Coastal Plain ancillariids may have evolved from one or more species of the Cretaceous-Paleocene genus Eoancilla. We agree with previous authors who have suggested that the late Eocene species Oliva mississippiensis Conrad, 1848 is the earliest known representative of this genus and the subfamily Oliviinae, perhaps derived from a species of Agaronia. The oldest Agaronia is Lower Eocene (Ypresian).


These data were generated to investigate the evolutionary relationships of olivoid gastropods living in the Paleocene and Eocene of the U.S. Gulf Coastal Plain. Nineteen species, representing three genera of Olividae and seven of Ancillaridae, were included along with Recent species Agaronia testacea (Lamarck, 1811) and Oliva sericea (Roding, 1798). Calipers were used to measure and code these specimens for 27 discrete characters: 10 binary, and 17 multistate. Character suites were coded to capture variation among (1) suture; (2) callus; (3) bands (including the olivoid, anterior, subsutural, and body whorl bands); (4) columella and plications; (5) ornamentation and texture; and (6) shell shape. Eoancilla was designated as the outgroup because the genus is a putative ancestor of the other ancillariids. Specimen measurements were made with calipers (measurements tallied in Supplement 2) and these coded character states were run in parsimony phylognetic analysis (detailed in Supplement 2). A parsimony analysis was conducted in PAUP* v. 4.0a147 (Swofford, 2003) using a heuristic search with 10,000 random addition sequences. TBR (tree bisection reconnection) was used for the branch-swapping algorithm with no reconnection limit, and collapsing all branches with a maximum branch length of zero. All characters were left unordered and equally weighted. Nexus files utilized are provided as Supplement 1. Values for the consistency index (CI) and retention index (RI) were recorded for the recovered trees, and bootstrap values and Bremer support were calculated using PAUP*. 

Usage notes

Supplement 1: Word file containing character matrix and Nexus files created with PAUP v. 4.0a147; can be opened with Microsoft Word or Google Docs. PAUP*, a free program, can be used to run the character matrix under similar phylogenetic analyses. The code and parameters of the phylogenetic analysis are included in the Word file, alongside the character matrix that was used.

Supplment 2: Spreadsheet of measurements can be opened using Excel or Google Sheets.


National Science Foundation, Award: NSF-EAR 0719642