xread 'Morphological matrix of extant and fossil Juglandaceae, modified from combined matrix of Manos et al. (2007), TreeBASE Study ID S1715, TreeBASE Legacy Study ID S1678. Paleooreomunnea stoneana rescored-carpel alignment & number of chambers at base of fruit unknown. Alatonucula ignis added. Citations for matrix: Manos, PS, and DE Stone. 2001. Evolution, phylogeny, and systematics of the Juglandaceae. Annals of the Missouri Botanical Garden 88: 231-269. Manos, PS, PS Soltis, DE Soltis, SR Manchester, S-H Oh, CD Bell, DL Dilcher, and DE Stone. 2007. Phylogeny of extant and fossil Juglandaceae inferred from the integration of molecular and morphological data sets. Systematic Biology 56: 412-430.'64 34 Rhoiptelia_chiliantha 00000000000000??00?????000001000000100????000100000010100000?000 Alfaroa_costaricensis 100111[01]11111000011100000010010110001111100530101202010101010101? Alfaroa_guanacastensis 1001111111110000111000000100101100011111005301012020101?1010101? Alfaroa_manningii 1001111111110000111000000100101100011111005301012020111?1010101? Alfaroa_williamsii 1001111111110000111000000100101100011111005301012020101?1010101? Alfaropsis_roxburghiana 1001011010110000011000100000001100121110001200001020111?00100012 Carya_cathayensis 0001000000021112110110210101201111231111137211114001?011011102?? Carya_cordiformis 01010000000211121101102101012011112311111372111140010011001102?? Carya_glabra 01010000000211121101102101012011112311111372111140010011001002?? Carya_illinoinensis 01010000000211121101102101012011112311111372111140010001001102?? Carya_myristiciformis 01010000000211121101102101012011112311111372111140010011001002?? Carya_ovata 01010000000211121101102101012011112311111372111140010011001102?? Carya_sinensis 110100100012111111?100210101201110231111137011112?01101?000012?? Carya_tomentosa 01010000000211121101102101012011112311111372111140010011001002?? Carya_tonkinensis 000100000002111211011021010120111123111113721111400110110?1?02?? Cruciptera_simsonii ???????0?????????1?????0??????11?0?011??1??21?????1?????0?010021 Cyclocarya_paliurus 001100000002101201011000[01]100201100201111142[02]11??0210101101000221 Engelhardia_serrata 1001010010110000011000?001001?110020111001120000?020100100100012 Engelhardia_spicata [01]00101[01]010110000011000000100111100201110011200001020001?00100012 Juglans_cinerea 01110000000210131111100211012011000011111260111030100001010101?? Juglans_mandshurica 01110000000210131111100211012011000011111260111030100001010102?? Juglans_microcarpa 01110000000210131101100211012011000011111262111030120001010102?? Juglans_nigra 01110000000210131101100211012011000011111262111030120001010102?? Juglans_regia 011100[01]0000210131111100211012011000011111262110030100011010102?? Oreomunnea_mexicana 100111[01]111110000111000000100101100011110011301012020111010101012 Oreomunnea_pterocarpa 1001111111110000111000?00100101100011110011301012020111?10101012 Paleooreomunnea_stoneana ?????001????????11?00??00??01011?0??1?10011?0??????????0101?0012 Paleoplatycarya_wingii 0?????0???????0?11?????00110101110011?001240??????0??1?010??0001 Platycarya_americana 0?????00??????0?1111?1200110101110011?0012401????????1?010?00001 Platycarya_strobilacea 0101000000021100111101200010001110011100124010000103001121100001 Polyptera_manningii 0???000????????2?101???000011011?0011?1?14?21???????011100?10021 Pterocarya_macroptera 0111000000121013010110001100201100001111123210000010100101010111 Pterocarya_stenoptera 00110[01]0000121013010110001100201100001111123210000010100101110111 Alatonucula_ignis ??????????????????????????????1120??1?10[01]012???????????????0001? ; cc - 0.63; proc /; optcode u 0.63; # $ ; cn {0 Tree_phenology strictly_deciduous evergreen; {1 Terminal_buds strictly_naked with_bud_scales; {2 Branchlet_pith solid chambered; {3 Stipules present absent; {4 Leaf_phyllotaxy alternate opposite; {5 Leaflet_arrangement odd-pinnate even-pinnate; {6 Leaflet_margin serrate strictly_entire; {7 Texture_of_lower_leaflet_surface smooth_to_verrucose papillose; {8 Druse_crystals_in_leaf_palisade present absent; {9 Rhomboid_crystals_in_leaf_palisade absent present; {10 Wood_porosity ring_and_semi-ring diffuse; {11 Vessel_perforation_plates exclusively_scalariform scalariform_in_part strictly_simple; {12 Mean_vessel_element_length >650_microns <650_microns; {13 Vessel_pore_shape angular_to_round strictly_round; {14 Inflorescences androgynous strictly_unisexual,_with_pistillate_terminal_and_staminate_lateral; {15 Number_of_staminate_catkins_per_inflorescence variable_1-10 5-8 clusters_of_3-5 exclusively_single; {16 Pistillate_catkin_stance pendent strictly_erect_at_anthesis; {17 Flower_sexuality bisexual unisexual; {18 Staminate_flower pedicellate sessile; {19 Staminate_bract_lobation 3-lobed_distally 1-lobed_distally; {20 Staminate_bract_vasculature 3_or_more_vascular_strands single_strand; {21 Staminate_flower_bracteoles present absent; {22 Staminate_flower_sepals 4_present 2_present absent; {23 Pollen_pore_distribution equatorial subequatorial heteropolar; {24 Pollen_aperture_number predominately_triporate 4_or_more; {25 Pollen_aperture_shape meridionally_elongate oblong_to_circular; {26 Pollen_sculpturing pseudocolpi_absent bipolar_pseudocolpi_present; {27 Pollen_wall exine_not_thinning_in_the_proximal_pole_area exine_thinning_in_the_area_surrounding_the_proximal_pole; {28 Pollen_size very_small,_ca_15_microns 17-26(-30)_microns,_rarely_larger >26_microns; {29 Pollen_wall_structure nexine_moderately_thick,_equal_to_ca_one-third_thickness_of_ektosexine nexine_extremely_thin,_less_than_one-third_thickness_of_ektosexine; {30 Ovary_position superior inferior; {31 Ovary 2-loculed 1-loculed; {32 Carpel_alignment median transverse oblique; {33 Stigmatic_disk absent present; {34 Stigma_orientation carinal split-carinal commissural; {35 Stylar_morphology two_long_style_arms two_short_style_arms four-lobed no_style_arms,_only_disk; {36 Ovule_type anatropous orthotropous; {37 Ovule_integument bitegmic unitegmic; {38 Pistillate_bract_in_fruit free attached; {39 Pistillate_bract_in_fruit enlarged small_or_fused_to_other_parts; {40 Pistillate_bract_lobation trilobed unlobed; {41 Pistillate_bracteole_configuration bracteoles_rudimentary_or_absent,_ovary_and_stigmas_exposed bracteoles_2,_posterior,_obscuring_the_ovary_but_stigmas_exposed bracteoles_2,_posterior,_obscuring_the_ovary_and_stigmas bracteoles_2,_lateral,_fused_to_ovary_about_halfway_up bracteoles_2,_lateral,_fused_to_top_of_ovary bracteoles_3_to_5,_lateral_and_posterior,_fused_to_top_of_ovary; {42 Fruit_type simple_fruit accessory_fruit_with_wings_formed_by_unlobed_or_trilobed_bract wings_formed_by_bract_and_2_bracteoles wings_formed_by_2_bracteoles wings_formed_by_2_lateral_sepals_and_2_bracteoles accessory_fruit_without_wings-husk_formed_by_4_sepals husk_formed_by_bract,_2_bracteoles,_and_4_sepals husk_formed_by_bract_and_bracteoles; {43 Fruit_chambers_at_base 2 no_character_state 4 8; {44 Nutshell_sclerenchyma principally_fibers principally_isodiametric_sclereids; {45 Cotyledons epigeous hypogeous; {46 Cotyledonary_node one_gap two_or_more_gaps; {47 Cotyledonary_node two_traces three_or_more_traces; {48 Seed_oils linolenic>linoleic/palmitic_low linolenic>linoleic/palmitic_high palmitic_high/linoleic>oleic linoleic>oleic/palmitic_low oleic>linoleic/palmitic_low; {49 Base_chromosome_number x=16 x=15 x=28; {50 Stigmatic_area outer-facing_part_of_the_style_arms inner-facing_part_of_the_style_arms all_surfaces_of_the_style_arms; {51 Wood_idioblasts rhomboidal_crystals_absent present_in_unchambered_axial_parenchyma present_in_chambered_axial_parenchyma present_in_ray_parenchyma; {52 Lateral_buds compact elongate; {53 Lateral_leaflets sessile_or_subsessile petiolulate; {54 Mean_leaflet_number 12_or_more fewer_than_12; {55 Serration_spacing teeth_one_or_fewer_per_secondary_vein teeth_two_or_more_per_secondary_vein; {56 Intersecondary_veins_in_the_leaflets rare common; {57 Freely_ending_veinlets_of_the_higher_order_venation_on_leaves less_than_10_percent 10_percent_or_more; {58 Peltate_scales_on_the_leaflet less_than_100_microns_in_diameter greater_than_120_microns_in_diameter; {59 Nutshell_lacunae absent present; {60 Primary_septum_of_the_fruit thick thin; {61 Position_of_vascular_bundles_relative_to_ribs_on_nuts on_smooth,_unribbed_surface along_ridges follow_grooves; {62 Fruit_wing_orientation_relative_to_the_long_axis_of_the_nutlet parallel oblique perpendicular; {63 Fruit_wing_venation absent subparallel pinnate; ;