#NEXUS [ File output by Morphobank v3.0 (http://www.morphobank.org); 2018-06-27 18.32.19 ] BEGIN TAXA; DIMENSIONS NTAX=47; TAXLABELS 'Eudigraphis taiwanensis' 'Glomeris marginata' 'Cyliosoma sp.' 'Glomeridesmus sp.' 'Cowiedesmus eroticopodus' 'Archidesmus macnicoli' 'Petaserpes sp.' 'Brachycybe lecontii' 'Pseudopolydesmus sp.' 'Prostemmiulus sp.' 'Abacion magnum' 'Cleidogona sp.' 'Gaspestria genselorum' 'Narceus americanus' 'Cambala annulata' 'Cylindroiulus sp.' 'Crussolum spp.' 'Scutigerina weberi' 'Sphendononema guildingii' 'Scutigera coleoptrata' 'Lithobius forficatus' 'Eupolybothrus cavernicolus' 'Anopsobius giribeti' 'Paralamyctes validus' 'Devonobius delta' 'Craterostigmus tasmanianus' 'Craterostigmus crabilli' 'Mazoscolopendra richardsoni' 'Scolopendropsis bahiensis' 'Akymnopellis chilensis' 'Rhysida longipes' 'Alipes grandidieri' 'Cryptops hortensis' 'Theatops spinicaudus' 'Scolopocryptops sexspinosus' 'Newportia adisi' 'Mecistocephalus guildingii' 'Tygarrup javanicus' 'Notiphilides grandis' 'Himantarium gabrielis' 'Stenotaenia linearis' 'Henia brevis' 'Strigamia maritima' 'Hydroschendyla submarina' 'Kachinophilus pereirai' 'Hanseniella sp.' 'Scutigerella sp.' ; ENDBLOCK; BEGIN CHARACTERS; DIMENSIONS NCHAR=205; FORMAT DATATYPE=STANDARD MISSING=? GAP=- SYMBOLS="01234567"; CHARLABELS [1] 'Egg tooth on embryonic cuticle of second maxilla' [2] 'Pattern of segment addition in ontogeny' [3] 'Fully-formed trunk leg pairs in first post-embryonic/post-pupoid stadium' [4] 'Number of fully-formed trunk leg pairs in first post-embryonic/post-pupoid stadium' [5] 'Brood care' [6] 'Peripatoid and foetoid stadia guarded by mother' [7] 'Antenna' [8] 'Antennal shape' [9] 'Variability in number of antennal articles' [10] 'Number of antennal articles' [11] 'Proportions of antennal articles' [12] 'Antennal trochanter' [13] 'Antennae with apical cones' [14] 'Schaftorgan on antennal scape' [15] 'Two lateral areas bearing sensilla basiconica on terminal antennal article' [16] 'Flattened head capsule' [17] 'Transverse cephalic suture' [18] 'Fenestrated plate composed of fused transverse tendons of mandibular, mx1, and mx2 segments' [19] 'Lateral eye' [20] 'Structure of lateral eye' [21] 'Four ocelli in rhomboid cluster' [22] 'Number of cellular components in eye' [23] 'Circumretinular sheath cells' [24] 'Interocellar sheath cells' [25] 'Tömösváry organs' [26] 'Mandibular glands' [27] 'Second vesicular gland' [28] 'Cuticle calcification' [29] 'Dentition of labral midpiece / intermediate part' [30] 'Labral sidepieces / lateral parts divided into a pair of alae by a transverse thickened line' [31] 'Single transverse seta projecting medially from labral side piece / lateral part' [32] 'Labral sidepiece / lateral part incised medially' [33] 'Lateral part of labrum bearing numerous strong, medially directed teeth' [34] 'A-shaped epipharyngeal support (labral trapezoid)' [35] 'Arching of border between labral and clypeal part of epipharynx' [36] 'Bilobate border between labral and clypeal parts of epipharynx' [37] 'Single row of bottle-shaped glandular openings at border between labral and clypeal parts of epipharynx' [38] 'Single row of bullet-shaped sensilla at proximal margin of field of branching spines at border between labral and clypeal parts of epipharynx' [39] 'Sensillar field(s) on medial clypeal part of epipharynx' [40] 'Elongate, figure-eight shaped group of two smooth scales surrounding each sensillum on clypeal part of epipharynx' [41] 'Lateral cluster of spear-shaped sensilla with inflated bases on clypeal part of epiphrynx' [42] 'Spear-shaped sensilla on proximal part of hypopharynx' [43] 'Stilus' [44] 'Four sclerites of mandible intersect at cruciform suture' [45] 'Mandible composed of two sclerites (lamina condylifera only sclerite differentiated from flank of mandible)' [46] 'Wide membranous band between sclerites of mandible, Haarpolster on a discrete sclerite' [47] 'Pectinate lamellae on mandible' [48] 'Arrangement of pectinate lamellae on mandible' [49] 'Dentate lamellae on mandible' [50] 'Number of groups of teeth in dentate lamella of mandible' [51] 'Internal and external teeth on mandible' [52] 'Field of fan bristles expanded onto lamina dentifera of mandible' [53] 'Molar plate' [54] 'Molar cusp on posterior part of molar plate' [55] 'First maxilla coalesced with sternal intermaxillary plate' [56] 'Median suture on first maxillary coxosternite' [57] 'Number of articles in telopodite of first maxilla' [58] 'Lappet on basal article of first maxillary telopodite' [59] '"Curled appendages" along inner margin of telopodite of first maxilla' [60] 'Brush-like setae along inner margin of telopodite of first maxilla' [61] 'Dense pad of setae on coxal process and inner margin of telopodite of first maxilla' [62] 'Distal part of telopodite and coxal process of first maxilla hyaline' [63] 'Lappets at base of coxal process of first maxilla' [64] 'Plumose setae on coxal process of first maxilla' [65] 'Maxillary organ' [66] 'Maxillary nephridia' [67] 'Limbless postmaxillary segment' [68] 'Second maxilla' [69] 'Coxae of second maxilla' [70] 'Metameric pores' [71] 'Metameric pores on second maxillary coxosternum' [72] 'Form of second maxillary telopodite' [73] 'Trochanter on second maxilla' [74] 'Pair of spine bristles at distal end of tibia of second maxilla' [75] 'Plumose setae on inner surface of tarsus of second maxillary telopodite' [76] 'Comb-like fringe of setae on distal article of telopodite of second maxilla' [77] 'Termination of telopodite of second maxilla' [78] 'Structure of claw of telopodite of second maxilla' [79] 'Forcipule with fang and venom gland' [80] 'Pleurite of forcipular segment arching over coxosternite' [81] 'Chitinous lines on forcipule' [82] 'Forcipular tooth plate' [83] 'Porodont on forcipular coxosternite' [84] 'Forcipular coxosternite sclerotised in midline' [85] 'Forcipular coxosternite deeply embedded into cuticle above second trunk segment' [86] 'Coxalplatten (forcipular coxosternal apodemes)' [87] 'Tarsungulum on forcipular/first trunk segment' [88] 'Spine comb on tarsus of forcipule/first trunk leg' [89] 'Basal node on forcipular tarsus/tarsungulum' [90] 'Basal node on forcipular femur' [91] 'Basal node on forcipular trochanteroprefemur' [92] 'Hinge between articles of forcipular telopodite' [93] 'Spine bristle on inner edge of forcipular trochanteroprefemur opposed to four spine bristles on each coxal margin' [94] 'Tergite of forcipular segment' [95] 'Width of forcipular tergite' [96] 'Number of post-cephalic leg-bearing segments' [97] 'Intraspecific variability in number of leg pairs' [98] 'Body segments fused into diplosegments' [99] 'Fusion of tergites, pleurites and sternites' [100] '"Special heterotergy" (alternating long and short tergites, with reversal of lengths between seventh and eighth walking leg-bearing segments)' [101] 'Overlap between tergite 1 and head shield' [102] 'Second tergite much larger than the following ones' [103] 'Single large tergal plate over trunk segments 7-9' [104] 'Long tergites divided into two' [105] 'Intercalary sclerites' [106] 'Principal paratergites' [107] 'Intercalary paratergites' [108] 'Paramedian sutures or grooves on tergum (Längsnähten)' [109] 'Crescentic sulci on most trunk tergites' [110] 'Tergite margination' [111] 'Tergite projections' [112] 'Large unpaired tergal spines associated with spine bristles, aligned longitudinally on midline' [113] 'Tergal spicula (hairs)' [114] 'Ultimate tergite with longitudinal median suture' [115] 'Anterior trunk sternites with rhachis (apodeme corresponding to mid-longitudinal sulcus)' [116] 'Complete sternal paramedian sutures along length of trunk' [117] 'Paired sternal paramedian pores with valves' [118] 'Sternal pore areas/sternal glands' [119] 'Suture in cuticular ring around sternal pores' [120] 'Definition of sternal pore area' [121] 'Endosternite' [122] 'Short, dense setae concentrated on posterior part of sternite of ultimate leg-bearing segment and inner part of ultimate leg coxa' [123] 'Leg pentagonal in cross-section, with marginal spines on the angles' [124] 'Proliferation of telopodal glands on posterior legs' [125] 'Socketed spurs D/V, a/m/p on distal extremities of podomeres' [126] 'Distal spinose projections on tibiae (tooth-like process on lateral side of at least tibiae 1-11)' [127] 'Bipartite division of tarsi of anterior series of trunk legs' [128] 'Tarsi divided into many joints' [129] 'Tarsal spurs' [130] 'Tarsus 1 with pair of terminal spurs' [131] 'Tarsal papillae and resilient sole hairs' [132] 'Coxal pouches on legs' [133] 'Ultimate leg-bearing segment a complete cylinder' [134] 'Coxopleurite' [135] 'Coxopleurites on ultimate legs' [136] 'Ultimate leg coxopleural process' [137] 'Ultimate leg thickened in males' [138] 'Heavily sclerotised, forcipulate ultimate legs' [139] 'Ultimate leg trochanter' [140] 'Trochanter of ultimate and penultimate legs with ventral spine' [141] 'Ultimate leg distomedial prefemoral process' [142] 'Ultimate leg pretarsus' [143] 'Antenna and leg regeneration' [144] 'Relationship of sternal and lateral longitudinal muscles' [145] 'Position of spiracle(s)' [146] 'Cribriform spiracles with floor of atrium raised into humps' [147] 'Spiracle with atrium divided by flapped valves' [148] 'stigmatic plates' [149] 'Shape of stigmatic plates' [150] 'First and second stigmatic plate' [151] 'Longitudinal and transverse connections between segmental tracheal branches' [152] 'Segmental arrangement of longitudinal connectives between spiracles' [153] 'Chiasmata' [154] 'Arrangement of tracheal pouches in comparison to body axis' [155] 'Apodemes of mid-body tracheal pouches' [156] 'Branching of mid-body tracheal pouch apodemes' [157] 'Foregut with differentiated gizzard with plicate walls' [158] 'Posteriorly directed spines along plicae of foregut' [159] 'Gizzard plicae covered by scales that each bear a single spine' [160] 'Posterior part of foregut organised as a sieve with stiff, anteriorly directed projections' [161] 'Anterior gizzard projections with pigmented bases bearing spinose scales or spines, distal part translucent, tapering, bearing dense hairs' [162] 'Shape of main sieve projections in gizzard' [163] 'Intestine shape' [164] 'Malpighian tubules formed as endodermal extensions of midgut' [165] 'Malpighian tubules formed as ectodermal extensions of hindgut' [166] 'Lateral defense glands' [167] 'Shape of defence glands' [168] 'Defence secretions with benzoquinines' [169] 'Unpaired median testis' [170] 'Testes differentiated into macrotestis with ampulla and microtestis' [171] 'Lateral testicular vesicles linked by a central, posteriorly extended deferens duct' [172] 'Testicular vesicles spindle shaped' [173] 'Number of testicular vesicles' [174] 'Female gonopod on first genital segment' [175] 'Female gonopod used to manipulate single eggs' [176] 'Female gonopod segmentation' [177] 'Female gonopod with basal article bearing spurs (macrosetae) and terminal article with a broad claw' [178] 'Claw of female gonopod fused with the apical article' [179] 'Position of male gonopore' [180] 'Male gonopod on second genital segment' [181] 'Form of male gonopod on second genital segment' [182] 'Bivalved anogenital capsule' [183] 'Anal organs' [184] 'Coxal organs' [185] 'Serial distribution of coxal organs' [186] 'Arrangement of coxal pores' [187] 'Spermatophore web produced by Spinngriffel on first genital segment of male' [188] 'Bean-shaped spermatophore with tough, multi-layered wall' [189] 'Ventral invagination in spermatophore' [190] 'Sperm dimorphism' [191] 'Sperm flagellum' [192] 'Coiling of sperm flagellum around nucleus' [193] 'Spiral ridge on nucleus of sperm' [194] 'Sperm nucleus with machette of microtubules' [195] 'Pseudoperforatorium' [196] 'Spermathecae formed by paired pockets in mouth cavity' [197] 'Modified leg(s) in male on seventh and/or eight trunk segment(s)' [198] 'Leg 8 in adult males' [199] 'Leg 9 in adult males' [200] 'Leg 10 in adult males' [201] 'Male gonopod development' [202] 'Cannula on gonopod coxa' [203] 'Posterior male leg pair as telopods with inner horns' [204] 'Terminal spinnerets on margin of epiproct, silk produced from setal sockets' [205] 'Pre-anal segment with trichobothria bearing a long sensory seta' ; STATELABELS 1 'absent' 'present' , 2 'hemianamorphosis' 'euanamorphosis' 'teloanamorphosis' 'epimorphosis' , 3 'Absent' 'Present' , 4 'three' 'four (Polyzoniida and Platydesmida coded as groundpatterns)' 'forcipule + four' 'forcipule + six' 'forcipule + seven' 'forcipule + eight' 'forcipule + 12' , 5 'absent' 'female (male in Brachycybe) bends ventrally around eggs' 'female bends dorsally around eggs' 'maternal brood sac on ventral side of opisthosoma' , 6 'absent' 'present' , 7 'Absent' 'Present' , 8 'filiform (antennal length > twice head width)' 'attenuate (antennal length < twice head width)' , 9 'variable within the species' 'fixed within the species' , 10 '15 or more' '14' 'eight or fewer' , 11 'at least some approximately as long as wide' 'consistently ring-like, much wider than long' , 12 'absent' 'present' , 13 'apical cones absent' 'apical cones present' , 14 'absent' 'present' , 15 'absent' 'present' , 16 'head capsule domed' 'flattened, with head bent posterior to the clypeus' , 17 'absent' 'present (frontal line or frontal sulcus)' 'present, divided near lateral margin into antenocellar suture' , 18 'absent' 'present' , 19 'Absent' 'Present' , 20 'stemmata' 'compound / facetted' 'single ocellus' , 21 'absent' 'present' , 22 'facultative constancy' 'all cellular components highly variable in number' , 23 'absent' 'present' , 24 'absent' 'present' , 25 'absent' 'present' , 26 'restricted to hypopharyngeal region' 'extended back into trunk' , 27 'absent' 'present' , 28 'absent' 'present' , 29 'undefined midpiece (median tooth / teeth lacking)' 'single strong tooth' 'three teeth' 'several (>3) small teeth' , 30 'side pieces undivided' 'alae definied by transverse thickened line' , 31 'absent' 'present' , 32 'not incised' 'incised' , 33 'absent' 'present' , 34 'absent' 'present' , 35 'subtransverse or gently arched distally' 'strongly arched distally' , 36 'absent' 'present' , 37 'absent' 'present' , 38 'absent' 'present' , 39 'medial cluster of sensilla proximally, near mouth opening' 'ovoid or crescentic fields of sensilla laterally' 'large field of sensilla across medial clypeal part of epipharynx, separated from spine field of a substantial expanse that bears scattered pores' 'band of sensilla medially, immediately proximal to spine field' 'lenticular field of sensilla immediately proximal to spine field' , 40 'absent' 'present' , 41 'absent' 'present' , 42 'absent' 'present' , 43 'absent' 'present' , 44 'absent' 'present' , 45 'absent' 'present' , 46 'absent' 'present' , 47 'Absent' 'Present' , 48 'multiple imbricated lamellae' 'lamellae arranged in a single file' , 49 'present' 'absent' , 50 'three' 'four/five' , 51 'absent' 'present' , 52 'absent' 'present' , 53 'Absent' 'Present' , 54 'absent' 'present' , 55 'absent' 'present, with unfused stipital and intermaxillary components' 'mental elements of gnathochilarium consolidated' , 56 'coxae medially coalesced, separated by median suture' 'coxae fused, without median suture' , 57 'two' 'one' , 58 'absent' 'present' , 59 'absent' 'present' , 60 'absent' 'with short, curved barbs' 'plumose, branching as slender hairs' , 61 'absent' 'present' , 62 'absent (telopodite sclerotised, non-hyaline, longer than and differentiated from coxal process)' 'present' , 63 'absent' 'present' , 64 'absent' 'present' , 65 'absent' 'present' , 66 'paired' 'fused' 'absent' , 67 'limbs present' 'limbs absent' , 68 'Absent' 'Present' , 69 'coxae separate' 'coxae fused' , 70 'Absent' 'Present' , 71 'minute opening of second maxillary gland medial to mx2 coxosternite' 'enlarged opening of mx2 gland ("metameric pore") incorporated in medial part of mx2 coxosternite' 'metameric pore on lateral part of mx2 coxosternite' , 72 'slender, leg-like, with elongate prefemur/femur' 'short, stout' , 73 'present' 'absent' , 74 'absent' 'present' , 75 'absent (simple setae)' 'plumose setae present' , 76 'absent' 'bifurcating of multifurcating spines' 'densely-aligned simple bristles' , 77 'simple (no claw or seta)' 'claw' 'seta or setigerous tubercle' , 78 'unipartite, conical claw' 'robust median claw with pair of slender spines on each side' 'thick, elongate digits with interspersed thin digits' 'fasciate claw' 'pectinate claw' 'hook-like claw' 'paired claw' , 79 'absent' 'present (fossils coded based on fang)' , 80 'absent (small pleurite)' 'pleurite arching over coxosternum, discontinuous medially ("Spange" of Attems, 1926)' 'pleurite arching over coxosternum, continuous ventromedially' , 81 'absent' 'present' , 82 'absent' 'anteriorly projecting serrate endite' 'transverse sclerotised band on anterior margin' , 83 'absent' 'translucent, seta-like porodont' , 84 'coxae separated medially, with sternite present in adult' 'coxosternal plates meeting medially, with flexible hinge' 'coxosternal plates meeting medially, hinge sclerotised and non-functional' , 85 'not embedded' 'deeply embedded' , 86 'absent' 'present' , 87 'separate tarsus and pretarsus' 'tarsus and pretarsus fused' , 88 'absent' 'present' , 89 'absent (coded for first trunk leg in Progoneata)' 'present' , 90 'absent (coded for first trunk leg in Progoneata)' 'present' , 91 'absent (coded for first trunk leg in Progoneata)' 'present' , 92 'between prefemur/trochanteroprefemur and femur' 'between trochanteroprefemur and tibia' 'between trochanteroprefemur and tarsungulum' , 93 'absent' 'present' , 94 'separate tergite' 'separate tergite lacking, fused to next posterior segment' , 95 'of similar width to head shield and T1' 'much narrower than head shield and T1, with forcipular pleurite strongly developed dorsally and coxopleural suture terminating dorsally' , 96 'forcipular segment + 15' 'forcipular segment + 21' 'forcipular segment + 23' 'forcipular segment + 37 or more' , 97 'constant number of leg pairs' 'variable number of leg pairs' , 98 'absent' 'present' , 99 'absent (free pleurites and sternites)' 'pleurotergites (pleurae fused to tergites, with free sternites)' 'complete body rings (tergites, pleurites and sternites fused)' , 100 'absent' 'present' , 101 'head shield overlaps tergite 1' 'tergite 1 overlaps head shield' , 102 'absent' 'present' , 103 'separate tergites' 'single tergite' , 104 'absent' 'present' , 105 'absent or weakly sclerotised' 'small intercalary tergites (pretergites) and sternites' 'strongly developed intercalary tergites and sternites' , 106 'absent' 'present' , 107 'absent' 'primary intercalary paratergites only' 'secondary intercalary paratergites present' , 108 'absent' 'present' , 109 'absent' 'present' , 110 'absent or on last tergite only' 'on most or all tergites' , 111 'absent on all tergites' 'present on at least TT11 and 13' , 112 'absent' 'present' , 113 'absent' 'present' , 114 'absent' 'present' , 115 'absent' 'present' , 116 'absent or incomplete' 'complete' , 117 'absent' 'present' , 118 'absent in at least female' 'present in both sexes' , 119 'absent' 'present' , 120 'sternal pore area on same level as surrounding parts of sternite' 'sternal pore area well-defined, circular or elongated, depressed relative to surrounding parts of sternite' , 121 'absent' 'present' , 122 'absent' 'present' , 123 'absent' 'present' , 124 'absent' 'present' , 125 'absent' 'present' , 126 'absent' 'present' , 127 'absent' 'present' , 128 'tarsi undivided or bisegmented' 'tarsus flagelliform, with many joints' , 129 'absent' 'present' , 130 'absent' 'present' , 131 'absent' 'present' , 132 'absent' 'present' , 133 'absent' 'present' , 134 'Absent' 'Present' , 135 'coxa and pleurites fused as short coxopleurite' 'elongate coxopleurite' , 136 'absent' 'present' , 137 'inconspicuous sexual dimorphism with respect to proportions of ultimate leg' 'present' , 138 'absent' 'present' , 139 'present' 'minute or absent' , 140 'absent' 'present' , 141 'absent' 'present' , 142 'claw' 'short, slender spine' 'tubercle' 'absent' , 143 'present' 'absent' , 144 'united sternal and lateral longitudinal muscles' 'separate sternal and lateral longitudinal muscles, with separate segmental tendons' , 145 'pleural' 'dorsal opening on tergum' 'sternal, at base of legs, with tracheal pouch serving as apodeme' 'single spiracle, opening on head' 'single spiracle on opisthomsomal segment 2' , 146 'absent' 'present' , 147 'valves absent' 'valves present' , 148 'Absent' 'Present' , 149 'flat, triangular' 'divided in midline' 'sterna nodifera sensu Blanke and Wesener (2014, character 20)' 'sterna costata sensu Blanke and Wesener (2014, character 20)' 'sliding sternites sensu Blanke and Wesener (2014, character 20)' , 150 'with tracheae' 'without tracheae, used for muscle attachment' , 151 'absent' 'present' , 152 'connected between successive spiracles' 'connected between spiracles separated by one or more segments' , 153 'absent' 'present' , 154 'diagonal orientation' 'longitudinal orientation' , 155 'absent' 'present, serving as msucle attachment' , 156 'uniramous' 'biramous' , 157 'absent' 'present' , 158 'absent' 'present' , 159 'absent' 'present' , 160 'absent' 'present' , 161 'absent' 'present' , 162 'evenly curved' 'kinked near midlength, with distal part more strongly directed forwards' , 163 'straight tube' 'N-shaped, bent twice' , 164 'absent' 'present' , 165 'absent' 'present' , 166 'absent' 'present' , 167 'elongate subtubular' 'subspherical' , 168 'absent' 'present (coded as grounpattern for Juliformia)' , 169 'symmetrically paired testis' 'one testis minute, undifferentiated' 'unpaired testis' , 170 'present' 'absent' , 171 'absent' 'present' , 172 'absent' 'present' , 173 'one pair' 'two or more pairs' , 174 'absent' 'present' , 175 'absent' 'present' , 176 'three articles and claw, with basal articles of gonopod pair separated' 'two articles, the proximal article of each gonopod pair partly joined (syntelopodite), the distal article a spine' 'single segment or with rudimentary second segment' , 177 'absent' 'present' , 178 'claw separate' 'claw fused' , 179 'opisthogoneate' 'behind coxa of second pair of trunk legs' 'through coxa of second pair of trunk legs' 'on fourth trunk segment' , 180 'present' 'absent' , 181 'blunt cones' 'slender styles' , 182 'absent' 'present' , 183 'absent' 'present through ontogeny' 'present only in juveniles' , 184 'absent' 'present' , 185 'on last four pairs of legs' 'on last two pairs of legs' 'on last pair of legs only' , 186 'few pores in linear row' 'numerous small pores scattered over coxopleure or large pore field' 'one or two large pores opening to expanded coxal organ' 'rosette of coxal organs opening into pit, without external pores' 'State 4' , 187 'absent' 'present' , 188 'absent' 'present' , 189 'absent' 'present' , 190 'absent' 'microsperm and macrosperm present' , 191 'present' 'absent' , 192 'absent (filiform)' 'present' , 193 'absent' 'present' , 194 'absent' 'present' , 195 'absent' 'present' , 196 'absent' 'present' , 197 'absent' 'present' , 198 'walking leg (unmodified)' 'accessory gonopod' 'functional gonopod' , 199 'walking leg (unmodified)' 'accessory gonopod' 'functional gonopod' , 200 'walking leg (unmodified)' 'accessory gonopod' 'functional gonopod' , 201 'gradual' 'abrupt' , 202 'absent' 'present' , 203 'absent' 'present' , 204 'absent' 'present' , 205 'absent' 'present' ; MATRIX 'Eudigraphis taiwanensis' 0010001?120010000?11-0001??0000000?????????000100?00101?-?------0?10----------0------?00000000--0100?00-0--00?00000000--00000010000100--00?00??02001000?000-0--???0010--?????0----1--000--0????????00000--000 'Glomeris marginata' 0010001?12001000001000001??11000?0?????????000100?1?102?--------0010----------0------?00000000--0100?10-0--00?00000000--00000000000000--?0??0??02001101?00110--???1010--?????0----1--000--00001-00000000--100 'Cyliosoma sp.' ?0??001?1200100000100???1??1100000?????????000100?10102?--------0?10----------0------?00000000--0100?10-0--00?00000000--00000000000000--?0?00???2001101?0011??????1010--?????0----1--000--0??01-00000000--100 'Glomeridesmus sp.' ?0??001?120010000?0-----1??1100??0?????????000100?10102?--------0?10----------0------?00000000--0100?00-0--00?00000000--00000000000100--?0??0???2?01101?0011??????1010--?????0----1--000--?????????00000--100 'Cowiedesmus eroticopodus' ??????1????0???0???????????1??????????????????????????????????????????????????0------????????0--?110?00-0--???00????1?????0?00?000?????????????????1????????????????????????????????????????????????1000????? 'Archidesmus macnicoli' ??????1???0????0??100??????1????????????????????????????--------??10----------0------??????000--?110?00-0--0??00????1?????0?000000???0--?0??????2??1?????????????????????????????????0??????????????1002??0?? 'Petaserpes sp.' ?111101?120010000?10????0??1000??0????????????????????2?--???---0?10----------0------?00000000--?100?00-0--00?00000000--00000000000100--?0??0???2?01210-0001??????001100?????0----1--000--0????????0101200000 'Brachycybe lecontii' ?111101?120010000?0-----0??1000??0??????????00100?1?102?--???---0?10----------0------?00000000--?110?00-0--00?00000000--00000000000100--?0??0???2?01210-0010??????00110??????0----1--000--0????????0101200000 'Pseudopolydesmus sp.' ?210001?12001000000-----1??1200?00?????????000100?1?102?--------0?10----------0------?00000000--0120?00-0--00?00000000--00000000000000--?0?00??02001410-00100--???001110?????0----2--000--0??01-0000120011010 'Prostemmiulus sp.' ?1??001?120110000?100???0??1200??0?????????000100?1?112?--???---0?10----------0------?00000000--?110?00-0--00?00000000--00000000000100--?0??0???2?01310-0111??????001110?????0----2--000--0????????012-011010 'Abacion magnum' ??10001?120110000?100???1??1200??0???????????010????112?--???---0?10----------0------?00000000--?110?00-0--00?00000000--00000000000100--?0??0???2?01310-0111??????001110?????0----2--000--0????????0120011010 'Cleidogona sp.' ?210001?120010000?100???1??1200??0???????????010????112?--???---0?10----------0------?00000000--0110?00-0--00?00000000--00000000000100--?0??0???2?01310-0111??????001110?????0----2--000--0????????0122010010 'Gaspestria genselorum' ??????1????????0???????????1???????????????????????????????????????0----------0------????????0--?120?00-0--00?00??????????0?00?00????????0???????????????????????????1??????????????-?????????????????????0?? 'Narceus americanus' 0110001?1200100000100???0??1200?00?????????000100?1?102?--------0010----------0------?00000000--1120?00-0--00?00000000--00000000000000--?0?00??02001410-01100--???001111?????0----1--000--0??01-0010112000000 'Cambala annulata' ?1??001?120010000?100???0??1200??0???????????010????102?--???---0?10----------0------?00000000--?120?00-0--00?0000000???00000000000000--?0??0???2?01410-0110??????001111??????----1-?000--0????????0121000000 'Cylindroiulus sp.' ?110001?1200100000100???0??1200?00?????????000100?1?102?--------0010----------0------?00000000--1120?00-0--00?00000000--00000000000000--?0?0???02001410-01100--???001111?????0----1--000--0??01-0010112000000 'Crussolum spp.' ??????1????????????????????0??????????????????????????????????????????????????1??0?00????0000?????????????????????????????1?0001000?????????????????????????????????????????????????????????????????????????? 'Scutigerina weberi' ?0??0010001001002111-???1??01000010000000000011000001000000100001?01010000000-100000000100001000000100100--00000000000--0010001101100100000003001000--0-0---??????0010--?????1110-000000--0????????00000--000 'Sphendononema guildingii' ?0??0010000001001111-???1??01000010000000000011000001000000100001?01010001000-100000000100001000000100100--00001100000--0010001101100100000003001000--0-0---??????0010--?????1110-001000--0????????00000--000 'Scutigera coleoptrata' 10120010001001001111-00010101000010000000000011000001000000100001101010001000-100000000100001000000100100--00001100000--0010001101100100000003001000--0-0---0--0--0010--000--1110-001000--00010010000000--000 'Lithobius forficatus' 10140010000000012010011111001011000010000000101001000-000002000101011100101012100011011000000000000100000--00110000000--0001101000000100000000000000--0-0---0--0--0010--210--1101101-0210010010010000000--000 'Eupolybothrus cavernicolus' ?0??001000000001201001111??01011000010000?00100-01000-00000200010?011100101012100011011000000000000100000--00110000000--0001101000000100000000000000--0-0---?--0--0010--???--1101101-021011????????00000--000 'Anopsobius giribeti' ?014001000000001200-----1??01010000010000000101001000-00000200000?011100101012120011011000000000000100000--00100000000--0000010000000101000000000000--0-0---??????0010--100--1101001-011101????????00000--000 'Paralamyctes validus' ?0??0010000000012012-???1??01010000010000000101001000-00000200000?011100101012120001011000000000000100000--00110000000--0000011000000100000000000000--0-0---0--0--0010--?????1101001-011001????????00000--000 'Devonobius delta' ??????10??0?00011?0-----0??0?0??0???????????????0?????0???????????011100??????11010211101000000??001?000?--000000??000--1?0?001000??????0???????0??0--?????????????????????????????????1????????????0???????? 'Craterostigmus tasmanianus' ?0161010100000012012-1101110300000??0???000010??00000-001000000002011100100112110102111000000000000100011--00000000000--0000000010001110000100?10000--0?0---0--0--0010--211010----01-11124100?0010000000--000 'Craterostigmus crabilli' ??????10100000012012-???1??0300000????????0010??00000-00100000000?011100100112110102111000000000000100011--00000000000--0000000010001110000100?10000--0?0---??????0010--?????0----01-111241????????00000--000 'Mazoscolopendra richardsoni' ??????10?00????1???????????0???????????????????????????????????????1??????????1???????10?00201010001?000??????00??????????0?0??0????0110?0??0????????????????????????????????????????0?121??????????0000--0?? 'Scolopendropsis bahiensis' ?30-??100000000100101???0??01000000001210001001001010-00001010000?0110-010021111010211100012010(1,2)100110001--10100010100--0000001010000110011010010010--100---1110??001?--?????0----01-001211????????00000--000 'Akymnopellis chilensis' ?30-1?100000000100101???0??01000000001210001001001010-00001010000?0110-0100211110102111000120101000100001--10100010100--0000001000000111001010010010--100---1110??0010--?????0----01-001211????????00000--000 'Rhysida longipes' ?30-11100000000100101???0??01000001001110001001001010-00001010000?0110-0100211110102111000120101000110001--10100000000--0000001010000111001000010100--100---1010--0010--211110----01-00121111??????00000--000 'Alipes grandidieri' ?30-11100000000100101???0??01000001001110001001001010-00001010000?0110-0100211110102111000120101000110001--10100000000--0000001010000111001003010100--100---1010--0010--?????0----01-001211????????00000--000 'Cryptops hortensis' 130-111010000001000-----0??01000000001400001001001000-0000100000020110-01002151102021110000101010001100020011000000000--1000000000000110001000010000--100---1001000010--?????0----01-00121110?0010000000--000 'Theatops spinicaudus' ?30-111010000001000-----0??01000000001300001001001000-00001000000?0110-01002161101021110001201010001000020010000010000--0000000010000110011000010000--100---1001000010--?????0----01-001211????????00000--000 'Scolopocryptops sexspinosus' ?30-111010000001000-----0??01000000001300001001001010-00001000000?0110-0100214110202111000120102000100001--10100000000--0000000010000111001000010000--100---1001110010--?????0----01-001211????????00000--000 'Newportia adisi' ?30-111010000001000-----0??01000000001300001001001000-00001000000?0110-01002141102021110000201020001000020011000000000--1000001000000111001003010000--100---1001110010--?????0----01-001211????????00000--000 'Mecistocephalus guildingii' ?30-??1011000011100-----01001100000000?0001000101--00-0010000100020111201000101100021110011200130000000-20010000001000--1100000000000110100001110000--101---0--0--0010--211101020-01-011211????????00000--000 'Tygarrup javanicus' ?30-??1011000011100-----0??0110000??????001000101--00-00100001000?0111201000101100021110001200130000000-20010000001000--1100000000000110000001110000--1?1---0--0--0010--?????1020-01-011211????????00000--000 'Notiphilides grandis' ?30-??1111000011000-----0??03000001000?0110000101--00-01100000000?0111211000131100021110000200031000000-21100000000001000000000000000110100003110000--111---0-????0010--?????1020-01-000--1????????00000--000 'Himantarium gabrielis' ?30-??1111000011100-----0??03000101000?0100000101--00-00110000000?0111111000101110021110000200031000000-21210000000001110000000000000110100003110000--111---??????0010--211101020-01-001211??00010000000--000 'Stenotaenia linearis' ?30-211011000011000-----0??03000001100?0000000111--00-01010000100?0111111000101110021110000200031000000-20010000000001000000000000000110000000110000--101---0--0--0010--?????1020-01-011211???0???000000--000 'Henia brevis' ?30-??1011000011000-----0??030000011??????0000111--00-01100000000?01111110002-1110021110000200031000000-2000000000000111000000000000011010000(0,3)110000--101---??????0010--?????1020-01-011211????????00000--000 'Strigamia maritima' 130-211011000011000-----01003000001100?0000000111--00-0100000000020111111000101100021110100200031000000-20000000000001000000000000000110100000110000--101---0--0--0010--211101020-01-011211??00010000000--000 'Hydroschendyla submarina' ?30-??1011000011000-----0??03000101000?0110000111--00-01010000100?0111111000141100021110000200031000000-20010000000001000000000000000110100003110000--101---0--0--0010???????1020-01-001221????????00000--000 'Kachinophilus pereirai' ??????10?100?0?10?0-----0??0????????????????????????0?????0000?0??011??1?000101110021?10101200031000000-200?0000000001?0000?000000000110000000?????0--????????????????????????????01-0?120??????????0000--000 'Hanseniella sp.' 00???010000000000?0-----1??0000000??????0??0?00-0?00100?--------0?00----------0------?00000000-?0000?00-?--00000000000--00000000000?00--00?00???3000--?-?---??????0010--?????0----3--000--000??????10000--001 'Scutigerella sp.' ?0???010000000000?0-----1??0000000??????0??0?00-0?00100?--------0?00----------0------?00000000-?0000?00-?--00000000000--00000000000?00--00?00???3000--?-?---??????0010--?????0----3--000--00010000010000--001 ; ENDBLOCK; BEGIN NOTES; [Character comments] TEXT CHARACTER=3 TEXT='ADDED CHARACTER--'; TEXT CHARACTER=4 TEXT='Inapplicable when fully formed leg pairs absent'; TEXT CHARACTER=7 TEXT='ADDED CHARACTER--constant when outgroup removed'; TEXT CHARACTER=8 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=9 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=10 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=11 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=12 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=13 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=14 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=15 TEXT='Inapplicable when no antenna'; TEXT CHARACTER=19 TEXT='ADDED CHARACTER--SPLIT STRUCTURE OF LATERAL EYE CHARACTER INTO TWO'; TEXT CHARACTER=20 TEXT='Inapplicable if lateral eye is absent'; TEXT CHARACTER=21 TEXT='Inapplicable if no stemmata on lateral eye'; TEXT CHARACTER=22 TEXT='Inapplicable if lateral eye absent'; TEXT CHARACTER=23 TEXT='Inapplicable if lateral eye absent'; TEXT CHARACTER=24 TEXT='Inapplicable if lateral eye absent'; TEXT CHARACTER=48 TEXT='Inapplicable when pectinate lamellae are absent'; TEXT CHARACTER=50 TEXT='Inapplicable if dentate lamellae are absent'; TEXT CHARACTER=51 TEXT='Inapplicable if dentate lamellae are absent'; TEXT CHARACTER=53 TEXT='ADDED CHARACTER'; TEXT CHARACTER=54 TEXT='Inapplicable if molar plate is absent'; TEXT CHARACTER=55 TEXT='Contingent on presence of first maxilla'; TEXT CHARACTER=57 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=58 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=59 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=60 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=61 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=62 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=63 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=64 TEXT='Inapplicable when first maxilla coalesced with sternal intermaxiliary'; TEXT CHARACTER=68 TEXT='ADDED CHARACTER'; TEXT CHARACTER=69 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=70 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=71 TEXT='Inapplicable when pores are absent'; TEXT CHARACTER=72 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=73 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=74 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=75 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=76 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=77 TEXT='Inapplicable when second maxilla absent'; TEXT CHARACTER=78 TEXT='Inapplicable when claw is absent'; TEXT CHARACTER=80 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=81 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=82 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=83 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=84 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=85 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=95 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=96 TEXT='Inapplicable when forcipule absent'; TEXT CHARACTER=104 TEXT='Inapplicable when special heterotergy absent (?)'; TEXT CHARACTER=106 TEXT='Inapplicable when intercalary sclerites are not strongly developed'; TEXT CHARACTER=107 TEXT='Inapplicable when intercalary sclerites are not strongly developed'; TEXT CHARACTER=119 TEXT='Inapplicable when sternal pores are absent'; TEXT CHARACTER=120 TEXT='Inapplicable when sternal pores are absent'; TEXT CHARACTER=134 TEXT='ADDED CHARACTER'; TEXT CHARACTER=135 TEXT='Inapplicable when coxopleurite is absent'; TEXT CHARACTER=136 TEXT='Inapplicable when coxopleurite is absent'; TEXT CHARACTER=145 TEXT='State 4 removed after outgroups removed'; TEXT CHARACTER=148 TEXT='ADDED CHARACTER'; TEXT CHARACTER=149 TEXT='Inapplicable when stigmatic plates absent'; TEXT CHARACTER=150 TEXT='Inapplicable when stigmatic plates absent'; TEXT CHARACTER=152 TEXT='Inapplicable when longitudinal connectives absent'; TEXT CHARACTER=154 TEXT='Inapplicable when stigmatic plates absent'; TEXT CHARACTER=155 TEXT='Inapplicable when stigmatic plates absent'; TEXT CHARACTER=156 TEXT='Inapplicable when stigmatic plates absent'; TEXT CHARACTER=158 TEXT='Inapplicable when foregut with differentiated gizzard absent'; TEXT CHARACTER=159 TEXT='Inapplicable when foregut with differentiated gizzard absent'; TEXT CHARACTER=161 TEXT='Inapplicable if anterior projections absent'; TEXT CHARACTER=162 TEXT='Inapplicable if anterior projections absent'; TEXT CHARACTER=167 TEXT='Inapplicable if defence glands absent'; TEXT CHARACTER=168 TEXT='Inapplicable if defence glands absent'; TEXT CHARACTER=172 TEXT='Inapplicable if testicular vesicles absent'; TEXT CHARACTER=173 TEXT='Inapplicable if testicular vesicles absent'; TEXT CHARACTER=175 TEXT='Inapplicable if female gonopod absent from first segment'; TEXT CHARACTER=176 TEXT='Inapplicable if female gonopod absent from first segment'; TEXT CHARACTER=177 TEXT='Inapplicable if female gonopod absent from first segment'; TEXT CHARACTER=178 TEXT='Inapplicable if claw is absent'; TEXT CHARACTER=179 TEXT='State 3 present in removed taxa'; TEXT CHARACTER=180 TEXT='Applicable only if gonopod is opisthogoneate'; TEXT CHARACTER=181 TEXT='Applicable only if gonopod is present on second segment'; TEXT CHARACTER=185 TEXT='Inapplicable if coxal organs absent'; TEXT CHARACTER=186 TEXT='Inapplicable if coxal organs absent'; TEXT CHARACTER=192 TEXT='Inapplicable if sperm flagellum absent'; TEXT CHARACTER=201 TEXT='Inapplicable if legs modified in seventh or eighth segment (?)'; TEXT CHARACTER=202 TEXT='Inapplicable if legs modified in seventh or eighth segment (?)'; [Attribute comments] TEXT TAXON=46 CHARACTER=48 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=47 CHARACTER=48 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=46 CHARACTER=151 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=47 CHARACTER=151 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=46 CHARACTER=153 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=47 CHARACTER=153 TEXT='Changed from ''-'' to ''?"'; TEXT TAXON=2 CHARACTER=168 TEXT='Changed from "0" to ''-'''; ENDBLOCK; BEGIN ASSUMPTIONS; TYPESET * UNTITLED = unord: 1 - 54 56 - 70 72 - 83 85 - 98 100 - 106 108 - 205, ord: 55 71 84 99 107; ENDBLOCK;