Data from: Spontaneous rhythmic and tool-assisted drumming across variable tempo and technique in a captive chimpanzee
Data files
May 05, 2026 version files 144.03 KB
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drumming_iois_clean.csv
26.66 KB
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drumming_iois.csv
28.10 KB
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drumming_onsets.csv
25.12 KB
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drumming_ratios_clean.csv
26.93 KB
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drumming_ratios.csv
29.34 KB
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isoc_ratio_counts.csv
1.69 KB
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README.md
6.19 KB
Abstract
In short, a captive chimpanzee named Toon drummed alongside vocalizations for several minutes, including across multiple pant hoot displays. We annotate this observation with precise onset timing, calculate inter-onset intervals, and analyze rhythmicity according to acoustic production mode and pant hoot display phase. Full methods and details can be found in the accompanying publication, whose abstract reads: Rhythmic drumming on percussive instruments is a common element of music across human cultures. Chimpanzees drum on the buttress roots of trees and on man-made objects, and often combine their drumming with long-distance pant-hoot vocalizations. Previous studies have suggested that chimpanzees select drumming substrates for their acoustic properties and that chimpanzee drumming shares core elements of human musicality (e.g., non-random timing, isochronous - metronome-like rhythm, and de-contextualized production in captivity). But to what extent can chimpanzees flexibly control their drumming rhythm across percussive media and techniques? Here, we report on a long-lasting drumming session by a captive chimpanzee named Toon, which was performed across multiple pant-hoot displays and employed diverse action forms, including drumstick-on-drum tool-composite use. We find flexible (isochronous) rhythm production across drumming implements, techniques, and tempi. Further, we describe tool composite transport and reuse, selectivity of percussive acoustic properties associated with different vocal elements, and variable use of the facial expression “play face” associated with faster and less variable rhythms, potentially indicating intrinsic enjoyment of such rhythms. Together, these findings provide evidence for key elements of human musicality in Toon’s drumming, supporting the hypothesis of shared evolutionary roots of human and chimpanzee drumming.
Dataset DOI: 10.5061/dryad.gtht76j21
Description of the data and file structure
This repository contains the video, code, and data accompanying the publication "Spontaneous rhythmic and tool-assisted drumming across variable tempo and technique in a captive chimpanzee". We here present all raw data, processing scripts, and analysis files.
Files and variables
Files
File: video1.mp4
Description: Video showing the beginning of the recording including the first drumming hits and vocalizations of the drumming event.
File: video2.mp4
Description: Video of Toon’s pant-hoot display showing the several elements reported here, including drumstick-on-drum tool set transport and use, integration of drumming across pant-hoot phases, variable use of percussive media by pant hoot phase, and variable expression of “play face”.
File: video4.mp4
Description: Video of the fourth drumming bout showing a short-long-long pattern.
File: video3.mp4
Description: Video of Toon showing the “play face” facial expression that was associated with faster drumming (shorter inter-hit intervals).
File: drumming_ratios.csv
Description: All drumming ratio data (pre-cleaning) throughout the observation - see processing scripts for full creation history.
File: drumming_ratios_clean.csv
Description: All drumming ratio data (cleaned) throughout the observation - see processing scripts for full creation history.
File: drumming_iois.csv
Description: All IOI data (pre-cleaning) throughout the observation - see processing scripts for full creation history.
File: drumming_iois_clean.csv
Description: All IOI data (cleaned) throughout the observation - see processing scripts for full creation history.
File: drumming_onsets.csv
Description: All drumming onset timing data (cleaned) throughout the observation - see processing scripts for full creation history.
File: isoc_ratio_counts.csv
Description: Cleaned ratio count data across the bout - see processing scripts for full creation history.
File: full-data-and-code.zip
Description: Full data and code. Note that the .zip file contains its own README file. The folder contains three subfolders (data [with raw and processed subfolders], illustrations, and plots). All scripts to convert raw to processed data and data into plots and illustrations are found in the home directory. Details of data format and behaviour coding scheme can be found in the main text.
Variables
Variables in drumming_iois.csv
Note that these are inter-onset intervals, so each row represents the time between two consecutive onsets. When e.g. surface_same is True, it means that the surface being used is the same for both hits/onsets. When limb is hands, it means that both hits used the hands, etc.
Categorical variables:
acoustic_bout- Bout identifierdrumming_bout- Drumming bout identifierdisplay_phase- Phase of displayplay_face- Was play face present or not? (boolean)percussive_medium- Type of percussive mediumpercussive_surface- Surface being used for the hitsurface_same- Whether surface is the same as previous strike (boolean)limb- Limb usedlimb_same- Whether limb is the same as previous strike (boolean)hands_or_feet- Was hand or foot used?tool_or_limb- Was a tool or a limb used?ioi_i- Inter-onset interval index
Continuous variables:
ioi- Inter-onset interval (milliseconds)
Variables in drumming_onsets.csv
Categorical variables:
display_phase- Phase of displayacoustic_bout- Bout identifierdrumming_bout- Drumming bout identifierplay_face- Was play face present or not? (boolean)percussive_medium- Type of percussive mediumpercussive_surface- Surface being used for the hitlimb- Limb usedhands_or_feet- Was hand or foot used?tool_or_limb- Was a tool or a limb used?
Continuous variables:
onset- Onset time (milliseconds)
Variables in drumming_ratios.csv and drumming_ratios_clean.csv
Categorical variables:
acoustic_bout- Bout identifierdrumming_bout- Drumming bout identifierdisplay_phase- Phase of displaypercussive_medium- Type of percussive mediumplay_face- Was play face present or not? (boolean)percussive_surface- Surface being used for the hitlimb- Limb usedhands_or_feet- Was hand or foot used? (boolean)tool_or_limb- Was a tool or a limb used? (boolean)ratio_i- Ratio indexlimbs_same- Whether limbs were the same across the 3 onsets used to calculate the ratio
Counts:
n_surfaces- Number of surfaces usedn_limbs- Number of limbs used
Continuous variables:
ratio- Ratio value. The scale is from 0 to 1, where 0.5 represents perfect isochrony.
Variables in isoc_ratio_counts.csv
Note that here each row represents three consecutive onsets.
Categorical variables:
drumming_bout- Drumming bout identifier
Counts:
on_isochronous- Count of on-isochronous observationsoff_isochronous- Count of off-isochronous observationsother- Count of observations that were neither in the isochronous bin nor the off-isochronous bintotal_obs- Total observations
Continuous variables:
on_isochronous_norm- Normalized on-isochronous count (normalized by total observations)off_isochronous_norm- Normalized off-isochronous count (normalized by total observations)other_norm- Normalized other count (normalized by total observations)isoc_prob- Isochrony probability (calculated as on-isochronous count divided by the sum of on-isochronous and off-isochronous counts)isoc_prob_normalized- Normalized isochrony probability (normalized by what is expected by chance assuming a uniform distribution of ratios; see Methods section of the paper for details)
