Data from: New information on Titanichthys (Placodermi: Arthrodira) from the Cleveland shale (upper devonian) of Ohio, USA
Data files
Aug 17, 2016 version files 62.27 KB
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Appendix 1.xlsx
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Appendix 2.docx
Abstract
The placoderm (Titanichthys) from the Late Devonian (Famennian) is based on incomplete and fragmentary specimens that have hindered understanding of its overall anatomy and phylogenetic relationships. A new, nearly complete, articulated specimen from the Upper Devonian Cleveland Shale provides new information about the previously undescribed rostral, postmarginal, postsuborbital, submarginal, posterior superognathal plates, and the nasal capsule. A revised diagnosis is provided for the genus. Three new diagnostic characters are identified including a transversely elliptical rostral plate that does not contact adjacent plates, a reduced posterior superognathal, and a median dorsal plate that inserts into the posterior dorsal lateral plate. The first comprehensive phylogenetic analysis of (Titanichthys) indicates that the genus is a basal aspinothoracid arthrodire closely related to the enigmatic taxa (Bungartius) and (Tafilalichthys).